This is a very interesting topic. Thanks for sharing the link!
I am reminded of a problem I had on my K8200. When the belts/pulleys were out of alignment, I would get stepping patterns in my prints, but when the pulleys were properly aligned, these artifacts would go away.
In theory, if the belts are tensioned and aligned properly then the number of teeth will be irrelevant. The belt should mesh with the teeth, and no positional errors should ever occur as a result of the teeth. Before masking the problem by optimizing the layer height, I would first make every effort to align and tension the belts. The better you get the machine adjusted, the better a baseline you will have for subsequent optimizations. (NOTE: Having the belts either too tight or too loose will cause problems.)
I am also reminded of a discussion in the K8200 forum about using DRV8255 stepper drivers, in which someone posted a diode fix for the stepper asymmetry. Here is the link to that discussion: